The Graf Lab @ The University of Alabama

email: dgraf(at) |
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dlgDr. Daniel L. Graf's
Curriculum Vitae

The following summarizes my career to-date. Use the following links to jump to various headings: Education, Professional Appointments, Grants, Awards & Fellowships, Service, Publications, Presentations & Posters, and Other Scholarly Products. More detailed information about my current research and teaching can be found elsewhere on this web site.

My CV on this web page was last updated 29 July 2013.


1997-2001 Ph.D. Biology, University of Michigan, Ann Arbor, Michigan.
Dissertation: A Phylogenetic Perspective on the Evolution of the Unionoida (Mollusca Bivalvia Palaeoheterodonta): Using Pattern to Test Hypotheses of Macroevolutionary Process.
1994-1997 M.Sc. Biology, Northeastern University, Boston, Massachusetts.
Thesis: Morphology, Zoogeography, and Taxonomy of Fusconaia flava (Rafinesque, 1820) (Mollusca: Bivalvia: Unionidae) in the Upper Mississippi, Great Lakes, and Nelson River Basins.
1989-1993 B.Sc. Biology, College of Biological Sciences, University of Minnesota, St. Paul, Minnesota.

Professional Appointments

2012-present Assistant Professor, Biology Department, University of Wisconsin-Stevens Point.
2008-present Research Associate, Academy of Natural Sciences of Philadelphia, Pennsylvania.
2004-present Research Associate, Illinois Natural History Survey, Champaign, Illinois.
2003-present Research Associate, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.
2008-2012 Assistant Professor and Full Member of the Graduate Faculty, Department of Biological Sciences, University of Alabama, Tuscaloosa, Alabama.
2002-2008 Assistant Curator, Malacology Department, Academy of Natural Sciences of Philadelphia, Pennsylvania.
2000-2001 Curatorial Assistant, Research Assistant and Collections Coordinator, Mollusk Division, University of Michigan Museum of Zoology, Ann Arbor, Michigan.
1995-1997 "Casual Help" and Curatorial Assistant, Malacology Department, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts.
1993-1994 Laboratory Assistant, Bell Museum of Natural History, University of Minnesota, St. Paul.

Recent Grants, Awards & Fellowships

2013-2014 ITIS, “Integrated Taxonomic Information System (ITIS) database development for freshwater mussels (Unionoida),” collaboration with Kevin Cummings (Illinois Natural History Survey): $13,000.
2013-2014 Undergraduate Education Initiative (UEI), UWSP, “Curation of the Daniel Bereza Collection of Freshwater Mussels at the National Museum of Natural History”: $3880.
2007-2013 National Science Foundation, "Phylogeny on the half-shell — Assembling the Bivalve Tree of Life." Senior Investigator with Gonzalo Giribet (Museum of Comparative Zoology), DEB-AToL-0732903, $137,574.
2007-2008 IUCN Freshwater Biodiversity Unit, Central Africa freshwater biodiversity assessment project, "Unionoida Assessment." 510 euros.
2006-2009 National Science Foundation, "Evolution in a Vortex: An Inventory of the Fishes and Mollusks of the Lower Congo River Rapids." Collaboration with Melanie Stiassny (American Museum of Natural History), DEB-BSI-0542575: $139,550 (including a supplemental award of $14,295).
2003-2008 National Science Foundation, "Systematic Revision of the Etherioidea, the Freshwater Mussels (Mollusca: Unionoida) of the Gondwanan Continents." Collaboration with Kevin Cummings (Illinois Natural History Survey), DEB-REVSYS-0316125: $199,805.

Recent Service

2015-2017 American Malacological Society, Council member at-large.
2009 IUCN Pan-Africa Freshwater Biodiversity Assessment Workshop participant and mollusk assessment evaluator, Cairo, Egypt, 3-7 May 2009.
2007-present Journal of Molluscan Studies, Associate Editor.
2007 American Malacological Society, Acting Chair of the Student Awards Committee during the 2007 World Congress of Malacology, Antwerp, Belgium, 16-20 July 2007.
2002-2004 American Malacological Society, Web Site Editor.
2002-2003 American Malacological Society, Systematics and Student Awards committees member.
2002-present NSF Proposal Reviews.
2000-present Numerous journal (and other scholarly publications) reviews.

Recent Publications

† indicates student collaborators. Click here for a complete list of my publications.

Combosch, D.J., T.M. Collins, E.A. Glover, D.L. Graf, E.M. Harper, J. Healy, G.Y. Kawauchi, S. Lemer, E. McIntyre, E. Strong, J.D. Taylor, J.D. Zardus, P.M. Mikkelsen, G. Giribet & R. Bieler. 2017. A family-level tree of life for bivalves based on a Sanger-sequencing approach. Molecular Phylogenetics & Evolution, in press. doi:10.1016/j.ympev.2016.11.003

† Pfeiffer, J.M. & D.L. Graf. 2015. Evolution of bilaterally asymmetrical larvae in freshwater mussels (Bivalvia: Unionoida: Unionidae). Zoological Journal of the Linnean Society 175: 307-318. doi:10.1111/zoj.12282

† Graf, D.L., H. Jones, A.J. Geneva, J.M. Pfeiffer & Michael W. Klunzinger. 2015. Molecular phylogenetic analysis supports a Gondwanan origin of the Hyriidae (Mollusca: Bivalvia: Unionida) and the paraphyly of Australasian taxa. Molecular Phylogenetics & Evolution 85: 1-9. doi:10.1016/j.ympev.2015.01.012

Cummings, K.S. & D.L. Graf. 2014. Mollusca: Bivalvia. pp. 423-506 in J.H. Thorp & D.C. Rogers (eds.). Ecology and General Biology: Thorp & Covich’s Freshwater Invertebrates. Academic Press-Elsevier, New York. ISBN: 978-0-12-385026-3

Graf, D.L., A.J. Geneva†, J.M. Pfeiffer III† & A.D. Chilala. 2014. Phylogenetic analysis of Prisodontopsis Tomlin, 1928 and Mweruella Haas, 1936 (Bivalvia: Unionidae) from Lake Mweru (Congo Basin) reveals endemic Quaternary radiation in the Zambian Congo. Journal of Molluscan Studies 80: 303-314. doi: 10.1093/mollus/eyu012

Recent Presentations & Posters

† indicates student collaborators. Click here for a complete list of presentations & posters.

Graf, D.L. 2017. Conflict Clade Analysis (CCA) of the freshwater mussel subfamily Unioninae (Bivalvia: Unionoida) [oral presentation]. American Malacological Society, 17-21 July 2017, University of Delaware, Newark, DE.

Cummings, K.S. & D.L. Graf. 2017. The freshwater mussels (Bivalvia: Unionida) of South America [poster]. Biennial Freshwater Mollusk Conservation Society Symposium, 26-30 March 2017, Cleveland, OH.

Cummings, K.S. & D.L. Graf. 2016. The freshwater mussels (Bivalvia: Unionoida) of South America and their conservation status [oral presentation]. Society for Freshwater Science Annual Meeting, 21-26 May 2016, Sacramento, CA.

Cummings, K.S. & D.L. Graf. 2015. The freshwater mussels (Bivalvia: Unionoida) of South America: a review [invited plenary speaker]. Second International Meeting on Biology and Conservation of Freshwater Bivalves, 4-8 October 2015, Buffalo, NY.

Graf, D.L., C. Jordan,† R. Pappas† & M. Zimbric.† 2015. Biogeography of the freshwater mussels (Unionida: Unionidae & Margaritiferidae) of Wisconsin, USA [oral presentation]. American Malacological Society, 28-31 August 2015, University of Michigan Biological Station, Pellston, MI.

Other Recent Scholarly Products

Graf, D.L. 2008. Book review of "Freshwater Mussel Ecology- A Multifactor Approach to Distribution and Abundance by David L. Strayer." The Malaclogist 51:!musselreview51.htm

Graf, D.L. & K.S. Cummings. 2002-2017. "The MUSSEL Project Web Site." Web site dedicated to explaining freshwater mussel diversity and evolution:

Graf, D.L. 2012-2017. "WInvertebrates: Towards an Account of the Freshwater Invertebrate taxa of Wisconsin, USA." Web site dedicated to synthesizing information of the diversity and zoology of the non-parasite, non-insect invertebrates of Wisconsin and adjacent areas:

Last updated 30 August 2017